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<h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Nebulidia</span></h1>
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<th>Nebulidae
</th></tr>




<tr>
<th><a href="Systematik_(Biologie)" title="Systematik (Biologie)">Systematik</a>
</th></tr>
<tr>
<td>
<table class="toptextcells" style="width:100%;">
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<td><i><a href="Reich_(Biologie)" title="Reich (Biologie)">Reich</a>:</i>
</td>
<td><a href="Disparia" title="Disparia">Disparia</a>
</td></tr>
<tr>
<td><i>ohne&nbsp;Rang:</i>
</td>
<td><a href="Provora" title="Provora">Provora</a>
</td></tr>
<tr>
<td><i><a href="Stamm_(Biologie)" title="Stamm (Biologie)">Stamm</a>:</i>
</td>
<td>Nebulidia
</td></tr>
<tr>
<td><i><a href="Klasse_(Biologie)" title="Klasse (Biologie)">Klasse</a>:</i>
</td>
<td>Nebulidea
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<td><i><a href="Ordnung_(Biologie)" title="Ordnung (Biologie)">Ordnung</a>:</i>
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<td>Nebulidida
</td></tr>


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<td><i><a href="Familie_(Biologie)" title="Familie (Biologie)">Familie</a>:</i>
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<td>Nebulidae
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<tr>
<th><a href="Nomenklatur_(Biologie)" title="Nomenklatur (Biologie)">Wissenschaftlicher Name</a> des&nbsp;<a href="Stamm_(Biologie)" title="Stamm (Biologie)">Stamms</a>
</th></tr>
<tr>
<td class="taxo-name">Nebulidia
</td></tr>
<tr>
<td class="Person">Tikhonenkov <i>et&nbsp;al.</i> 2022<sup id="cite_ref-Tikhonenkov2022-12_1-0" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
</td></tr>
<tr>
<th><a href="Nomenklatur_(Biologie)" title="Nomenklatur (Biologie)">Wissenschaftlicher Name</a> der&nbsp;<a href="Klasse_(Biologie)" title="Klasse (Biologie)">Klasse</a>
</th></tr>
<tr>
<td class="taxo-name">Nebulidea
</td></tr>
<tr>
<td class="Person">Tikhonenkov <i>et&nbsp;al.</i> 2022<sup id="cite_ref-Tikhonenkov2022-12_1-1" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
</td></tr>
<tr>
<th><a href="Nomenklatur_(Biologie)" title="Nomenklatur (Biologie)">Wissenschaftlicher Name</a> der&nbsp;<a href="Ordnung_(Biologie)" title="Ordnung (Biologie)">Ordnung</a>
</th></tr>
<tr>
<td class="taxo-name">Nebulidida
</td></tr>
<tr>
<td class="Person">Tikhonenkov <i>et&nbsp;al.</i> 2022<sup id="cite_ref-Tikhonenkov2022-12_1-2" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
</td></tr>
<tr>
<th><a href="Nomenklatur_(Biologie)" title="Nomenklatur (Biologie)">Wissenschaftlicher Name</a> der&nbsp;<a href="Familie_(Biologie)" title="Familie (Biologie)">Familie</a>
</th></tr>
<tr>
<td class="taxo-name">Nebulidae
</td></tr>
<tr>
<td class="Person">Tikhonenkov <i>et&nbsp;al.</i> 2022<sup id="cite_ref-Tikhonenkov2022-12_1-3" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
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<p><b>Nebulidia</b> ist ein <a href="Phylum" class="mw-redirect" title="Phylum">Phylum</a> (<a href="Stamm_(Biologie)" title="Stamm (Biologie)">Stamm</a>) räuberischer <a href="Mikrobiell" class="mw-redirect" title="Mikrobiell">mikrobieller</a> <a href="Eukaryoten" title="Eukaryoten">Eukaryoten</a> (<a href="Mikroeukaryoten" class="mw-redirect" title="Mikroeukaryoten">Mikroeukaryoten</a>, <a href="Protisten" title="Protisten">Protisten</a>) in der Supergruppe <a href="Provora" title="Provora">Provora</a>. Das Phylum dieser <a href="Einzeller" title="Einzeller">Einzeller</a> umfasst <a href="Monotypisch" title="Monotypisch">monotypisch</a> nur die <a href="Klasse_(Biologie)" title="Klasse (Biologie)">Klasse</a> <b>Nebulidea</b>, mit der einzigen <a href="Ordnung_(Biologie)" title="Ordnung (Biologie)">Ordnung</a> <b>Nebulidida</b>, und diese die eine <a href="Familie_(Biologie)" title="Familie (Biologie)">Familie</a> <b>Nebulidae</b> – diese Gruppe (<a href="Klade" title="Klade">Klade</a>) wird informell auch als ‚Nebuliden‘ (<span style="font-style:normal;font-weight:normal"><a href="Englische_Sprache" title="Englische Sprache">englisch</a></span> <span lang="en-Latn" style="font-style:italic">nebulids</span>) bezeichnet. Die Familie Nebulidae beinhaltet die <a href="Gattung_(Biologie)" title="Gattung (Biologie)">Gattungen</a> <i><a href="Nebulomonas" class="mw-redirect" title="Nebulomonas">Nebulomonas</a></i> und <i><a href="Ancoracysta" class="mw-redirect" title="Ancoracysta">Ancoracysta</a></i>.<sup id="cite_ref-NCBI_2-0" class="reference"><a href="#cite_note-NCBI-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Tikhonenkov2022-12_1-4" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Belyaev2024_3-0" class="reference"><a href="#cite_note-Belyaev2024-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup>
</p>

<div class="mw-heading mw-heading2"><h2 id="Nahrungserwerb">Nahrungserwerb</h2></div>
<p>Zum Nahrungserwerb besitzen die Nebulidia <a href="Organell#Taxonspezifische_Organellen" title="Organell">spezielle Organellen</a>, dei bei der Gattung <i><a href="Ancoracysta" class="mw-redirect" title="Ancoracysta">Ancoracysta</a></i> Ancoracysten und bei der Gattung <i><a href="Nebulomonas" class="mw-redirect" title="Nebulomonas">Nebulomonas</a></i> Amphoracysten (Toxicysten) genannt werden.<sup id="cite_ref-Tikhonenkov2022-12_1-5" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
</p>
<div class="mw-heading mw-heading2"><h2 id="Systematik">Systematik</h2></div>
<p>Die Systematik der Klade lautet mit Stand 9. Dezember 2025 wie folgt:
</p><p><a href="Reich_(Biologie)" title="Reich (Biologie)">Reich</a> <a href="Disparia" title="Disparia">Disparia</a> <small><style data-mw-deduplicate="TemplateStyles:r261921330">
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</style><span class="Person h-card">Valt &amp; Čepička</span> 2025</small><sup>(V)</sup> <span style="white-space:nowrap">[Promethea <small><span class="Person h-card">Zlatogursky, Boscaro &amp; Keeling</span>, 2025</small><sup id="cite_ref-Zlatogursky2025_4-0" class="reference"><a href="#cite_note-Zlatogursky2025-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup>]</span>
</p>
<ul><li><a href="Klade" title="Klade">Klade</a> <a href="Membrifera" title="Membrifera">Membrifera</a> <small><span class="Person h-card">Valt &amp; Čepička</span> 2025</small><sup>(V)</sup>
<ul><li><a href="Phylum" class="mw-redirect" title="Phylum">Phylum</a> <a href="Caelestes" title="Caelestes">Caelestes</a> <small><span class="Person h-card">Valt &amp; Čepička</span> 2025</small><sup>(V)</sup></li>
<li>Phylum <a href="Hemimastigophora" title="Hemimastigophora">Hemimastigophora</a> <small><span class="Person h-card"><a href="Wilhelm_Foissner" title="Wilhelm Foissner">W.&nbsp;Foissner</a>, H.&nbsp;Blatterer &amp; I.&nbsp;Foissner</span>, 1988</small></li></ul></li>
<li>Klade <a href="Provora" title="Provora">Provora</a> <small><span class="Person h-card">Tikhonenkov</span> <i>et&nbsp;al.</i> 2022</small><sup>(N,T,V)</sup>
<ul><li>Phylum <b>Nebulidia</b> <small><span class="Person h-card">Tikhonenkov</span> <i>et&nbsp;al.</i> 2022</small><sup>(N,T)</sup>
<ul><li>Klasse <b>Nebulidea</b><sup>(N)</sup> <small><span class="Person h-card">Tikhonenkov</span> <i>et&nbsp;al.</i> 2022</small><sup>(T)</sup>
<ul><li>Ordnung <b>Nebulidida</b><sup>(N)</sup> <small><span class="Person h-card">Tikhonenkov</span> <i>et&nbsp;al.</i> 2022</small><sup>(T)</sup>
<ul><li>Familie <b>Nebulidae</b><sup>(N)</sup> <small><span class="Person h-card">Tikhonenkov</span> <i>et&nbsp;al.</i> 2022</small><sup>(T)</sup>
<ul><li>Gattung <i><a href="Ancoracysta" class="mw-redirect" title="Ancoracysta">Ancoracysta</a></i> <small><span class="Person h-card">Janouškovec, Tikhonenkov, Burki, Howe, Rohwer, Mylnikov &amp; Keeling</span> 2017</small><sup>(N)</sup><sup id="cite_ref-Janouskovec2017_5-0" class="reference"><a href="#cite_note-Janouskovec2017-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li>
<li>Gattung <i><b><a href="Nebulomonas" class="mw-redirect" title="Nebulomonas">Nebulomonas</a></b></i> <small><span class="Person h-card">Tikhonenkov</span> <i>et&nbsp;al.</i> 2022</small><sup>(N,T)</sup></li></ul></li></ul></li></ul></li></ul></li>
<li>Phylum <a href="Nibbleridia" title="Nibbleridia">Nibbleridia</a> <small><span class="Person h-card">Tikhonenkov</span> <i>et&nbsp;al.</i> 2022</small><sup>(N,T)</sup><br>– mit Gattungen <i><a href="Nibbleromonas" title="Nibbleromonas">Nibbleromonas</a></i>, <i><a href="Ubysseya" class="mw-redirect" title="Ubysseya">Ubysseya</a></i></li></ul></li></ul>
<hr>
<dl><dd>(N) – <span lang="en"><a href="National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">National Center for Biotechnology Information</a></span> (NCBI)<sup id="cite_ref-NCBI_2-1" class="reference"><a href="#cite_note-NCBI-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></dd>
<dd>(V) – Marek Valt, Alexander K. Tice, Ivan Čepička <i>et&nbsp;al.</i> (2025)<sup id="cite_ref-Valt2025_6-0" class="reference"><a href="#cite_note-Valt2025-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup></dd>
<dd>(T) – Tikhonenkov <i>et&nbsp;al.</i><sup id="cite_ref-Tikhonenkov2022-03_7-0" class="reference"><a href="#cite_note-Tikhonenkov2022-03-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Tikhonenkov2022-12_1-6" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Tikhonenkov2023_8-0" class="reference"><a href="#cite_note-Tikhonenkov2023-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></dd></dl>
<div class="mw-heading mw-heading2"><h2 id="Phylogenie">Phylogenie</h2></div>
<p>Die phylogenetischen Beziehungen zwischen den beschriebenen Arten der Nebulidia werden wie folgt dargestellt:<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>1.1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>3.1<span class="cite-bracket">]</span></a></sup>
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<td style="min-width:1em; border-bottom:2px solid; vertical-align:bottom; text-align:center;">&nbsp;<a href="Provora" title="Provora">Provora</a><sup><sup><span style="visibility:hidden;">.</span></sup></sup>&nbsp;
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<td style="min-width:1em; border-bottom:2px solid; vertical-align:bottom; text-align:center;">&nbsp;<b>Nebulidia</b><sup><span style="visibility:hidden;">.</span></sup>&nbsp;
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<td style="min-width:1em; border-bottom:2px solid; vertical-align:bottom; text-align:center;">&nbsp;<i>Nebulomonas</i>&nbsp;
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<p><i><a href="Nebulomonas_marisrubri" title="Nebulomonas marisrubri">N. marisrubri</a></i>
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<td style="border-left:2px solid; border-bottom:2px solid; vertical-align:bottom; text-align:center;">&nbsp;<i>Ancoracysta</i>&nbsp;
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<p><i><a href="Ancoracysta_twista" title="Ancoracysta twista">A. twista</a></i>
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<td style="border-left:2px solid; border-bottom:2px solid; vertical-align:bottom; text-align:center;">&nbsp;<a href="Nibbleridia" title="Nibbleridia">Nibbleridia</a>&nbsp;
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<p><i><a href="Ubysseya_fretuma" title="Ubysseya fretuma">Ubysseya fretuma</a></i>, <i><a href="Nibbleromonas" title="Nibbleromonas">Nibbleromonas spp.</a></i>
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<td valign="top"><br><span style="display:none;">Vorlage:Klade/Wartung/Style</span>
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<div class="mw-heading mw-heading2"><h2 id="Etymologie">Etymologie</h2></div>
<p>Die Bezeichnungen leiten sich ab von <span style="font-style:normal;font-weight:normal"><a href="Latein" title="Latein">lateinisch</a></span> <span lang="la-Latn" style="font-style:italic">nebulosus</span>, deutsch <span lang="de" style="font-style:normal;font-weight:normal">‚neblig‘</span>, <a href="Englische_Sprache" title="Englische Sprache">englisch</a> <span lang="en" style="font-style:italic;font-weight:normal">misty</span>, der jeweilige Suffix kennzeichnet die taxonomische Rangstufe.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>1.2<span class="cite-bracket">]</span></a></sup>
</p>
<div class="mw-heading mw-heading2"><h2 id="Weiterführende_Literatur"><span id="Weiterf.C3.BChrende_Literatur"></span>Weiterführende Literatur</h2></div>
<ul><li>Sei Suzuki-Tellier, Federica Miano, Seyed Saeed Asadzadeh, Alastair G.&nbsp;B. Simpson, Thomas Kiørboe: <i>Foraging mechanisms in excavate flagellates shed light on the functional ecology of early eukaryotes.</i> In: <i><a href="PNAS" class="mw-redirect" title="PNAS">PNAS</a></i>, Band 121, Nr.&nbsp;22, 23. Mai 2024, S.&nbsp;e2317264121; <a href="https://doi.org/10.1073/pnas.2317264121" class="extiw external" title="doi:10.1073/pnas.2317264121">doi:10.1073/pnas.2317264121</a>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/38781211?dopt=Abstract">PMID 38781211</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11145212/">PMC&nbsp;11145212</a> (freier Volltext), <a href="ResearchGate" class="mw-redirect" title="ResearchGate">ResearchGate</a>:<a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/375822320">375822320</a> (englisch).</li>
<li>Łukasz Lamża: <i>Deep-branching eukaryotes and early events in protist evolution.</i> In: <i>Biological Reviews</i>, 9. November 2025; <a href="https://doi.org/10.1111/brv.70101" class="extiw external" title="doi:10.1111/brv.70101">doi:10.1111/brv.70101</a> (englisch).</li></ul>
<div class="mw-heading mw-heading2"><h2 id="Einzelnachweise">Einzelnachweise</h2></div>
<ol class="references">
<li id="cite_note-Tikhonenkov2022-12-1"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-Tikhonenkov2022-12_1-0">a</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-1">b</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-2">c</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-3">d</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-4">e</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-5">f</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-6">g</a></sup></span> <span class="reference-text">
Denis Victorovich Tikhonenkov, Kirill V. Mikhailov, Ryan M.&nbsp;R. Gawryluk, Artem O. Belyaev, Varsha Mathur, Sergey A. Karpov, Dmitry G. Zagumyonnyi, Anastasia S. Borodina, Kristina I. Prokina, Alexander P. Mylnikov, Vladimir V. Aleoshin, Patrick J. Keeling: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Microbial predators form a new supergroup of eukaryotes</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic"><a href="Nature" title="Nature">Nature</a></cite>. 612. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em">&nbsp;</span>7941</span>, 7.&nbsp;Dezember 2022, <a href="Internationale_Standardnummer_f%C3%BCr_fortlaufende_Sammelwerke" title="Internationale Standardnummer für fortlaufende Sammelwerke">ISSN</a>&nbsp;<span style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://zdb-katalog.de/list.xhtml?t=iss%3D%220028-0836%22&amp;key=cql">0028-0836</a></span>, <a href="ResearchGate" class="mw-redirect" title="ResearchGate">ResearchGate</a>:<a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/366093658">366093658</a>, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em">&nbsp;</span>714–719</span>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1038/s41586-022-05511-5">10.1038/s41586-022-05511-5</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/36477531?dopt=Abstract">PMID 36477531</a>, <a href="Bibcode" title="Bibcode">bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2022Natur.612..714T">2022Natur.612..714T</a> (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rfr_id=info:sid/de.wikipedia.org:Nebulidia&amp;rft.atitle=Microbial+predators+form+a+new+supergroup+of+eukaryotes&amp;rft.au=Denis+Victorovich+Tikhonenkov%2C+Kirill+V.+Mikhailov%2C+Ryan+M.+R.+Gawryluk%2C+...&amp;rft.date=2022-12-07&amp;rft.doi=10.1038%2Fs41586-022-05511-5&amp;rft.genre=journal&amp;rft.issn=0028-0836&amp;rft.issue=7941&amp;rft.jtitle=Nature&amp;rft.pages=714-719&amp;rft.pmid=36477531&amp;rft.volume=612.+Jahrgang" style="display:none">&nbsp;</span> Dazu:
<dl><dd><dl><dd><ul><li><a rel="nofollow" class="external text" href="https://static-content.springer.com/esm/art%3A10.1038%2Fs41586-022-05511-5/MediaObjects/41586_2022_5511_MOESM1_ESM.pdf">Supplementay Information</a> (PDF; 0,5&nbsp;MB)</li>
<li>Nadja Podbregar: <a rel="nofollow" class="external text" href="https://www.scinexx.de/news/biowissen/neue-supergruppe-am-baum-des-lebens/">Neue Supergruppe am Baum des Lebens</a> – Uralte Gruppe räuberischer Einzeller unterscheidet sich von allen anderen Lebewesen. Auf: <a href="Scinexx.de" class="mw-redirect" title="Scinexx.de">scinexx.de</a> vom 15. Dezember 2022.</li>
<li><a rel="nofollow" class="external text" href="https://www.sci.news/biology/provora-11460.html">Meet Provora, New Supergroup of Eukaryotes</a>. Auf <a href="Sci.News" title="Sci.News">Sci.News</a> vom 8. Dezember 2022.</li>
<li>Stefan Labbé: <a rel="nofollow" class="external text" href="https://www.vancouverisawesome.com/highlights/nibblers-and-lions-bc-researchers-discover-new-supergroup-on-tree-of-life-6220625">'Nibblers and Lions': B.C. researchers discover new 'supergroup' on tree of life</a>. Auf: Vancouver is Awesome vom 8. Dezember 2022.</li></ul></dd></dl></dd></dl>
<ol class="mw-subreference-list"><li id="cite_note-9"><span class="mw-cite-backlink"><a href="#cite_ref-9">↑</a></span> <span class="reference-text">S. 717</span>
</li>
<li id="cite_note-11"><span class="mw-cite-backlink"><a href="#cite_ref-11">↑</a></span> <span class="reference-text">Suppl. Inf.</span>
</li>
</ol></span></li>
<li id="cite_note-NCBI-2"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-NCBI_2-0">a</a></sup> <sup><a href="#cite_ref-NCBI_2-1">b</a></sup></span> <span class="reference-text">
<a href="National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a> Taxonomy Browser: <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?command=show&amp;mode=tree&amp;id=3004206&amp;lvl=3">Provora</a>, Details: <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?command=show&amp;mode=node&amp;id=3004206">Provora in [Tikhonenkov DV et al. (2022)]</a>. Graphisch: <a rel="nofollow" class="external text" href="https://lifemap.cnrs.fr/tree?efficiency-mode=true&amp;tool=search&amp;tid=3004206">Provora</a>, auf <a href="Lifemap" class="mw-redirect" title="Lifemap">Lifemap</a>.</span>
</li>
<li id="cite_note-Belyaev2024-3"><span class="mw-cite-backlink"><a href="#cite_ref-Belyaev2024_3-0">↑</a></span> <span class="reference-text">
Artem O. Belyaev, Sergei A. Karpov, Patrick J. Keeling, Denis V. Tikhonenkov: <cite class="lang" lang="en" dir="auto" style="font-style:italic">The nature of 'jaws': a new predatory representative of Provora and the ultrastructure of nibbling protists</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Open Biology</cite>. 14. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em">&nbsp;</span>12</span>, 18.&nbsp;Dezember 2024, <a href="ResearchGate" class="mw-redirect" title="ResearchGate">ResearchGate</a>:<a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/387173990">387173990</a>, <a rel="nofollow" class="external text" href="https://www3.botany.ubc.ca/keeling/PDF/24jaws.pdf">PDF</a> (botany.ubc.ca), <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em">&nbsp;</span>240158</span>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1098/rsob.240158">10.1098/rsob.240158</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/39689855?dopt=Abstract">PMID 39689855</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11651884/">PMC&nbsp;11651884</a> (freier Volltext) – (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rfr_id=info:sid/de.wikipedia.org:Nebulidia&amp;rft.atitle=The+nature+of+%27jaws%27%3A+a+new+predatory+representative+of+Provora+and+the+ultrastructure+of+nibbling+protists&amp;rft.au=Artem+O.+Belyaev%2C+Sergei+A.+Karpov%2C+Patrick+J.+Keeling%2C+...&amp;rft.date=2024-12-18&amp;rft.doi=10.1098%2Frsob.240158&amp;rft.genre=journal&amp;rft.issue=12&amp;rft.jtitle=Open+Biology&amp;rft.pages=240158&amp;rft.pmc=11651884&amp;rft.pmid=39689855&amp;rft.volume=14.+Jahrgang" style="display:none">&nbsp;</span></span>
<ol class="mw-subreference-list"><li id="cite_note-10"><span class="mw-cite-backlink"><a href="#cite_ref-10">↑</a></span> <span class="reference-text">S. 11-12</span>
</li>
</ol></li>
<li id="cite_note-Zlatogursky2025-4"><span class="mw-cite-backlink"><a href="#cite_ref-Zlatogursky2025_4-0">↑</a></span> <span class="reference-text">
Vasily Zlatogursky, Vittorio Boscaro, Gordon Lax, Matias Wanntorp, Nina Pohl, Fabien Burki, Patrick J. Keeling: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Phylogenetic position and mitochondrial genome evolution of “orphan” eukaryotic lineages</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic">iScience</cite>. 28. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em">&nbsp;</span>8</span>, August 2025, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em">&nbsp;</span>113184</span>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1016/j.isci.2025.113184">10.1016/j.isci.2025.113184</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/40948565?dopt=Abstract">PMID 40948565</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC12432456/">PMC&nbsp;12432456</a> (freier Volltext) – (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rfr_id=info:sid/de.wikipedia.org:Nebulidia&amp;rft.atitle=Phylogenetic+position+and+mitochondrial+genome+evolution+of+%E2%80%9Corphan%E2%80%9D+eukaryotic+lineages&amp;rft.au=Vasily+Zlatogursky%2C+Vittorio+Boscaro%2C+Gordon+Lax%2C+...&amp;rft.date=2025-08&amp;rft.doi=10.1016%2Fj.isci.2025.113184&amp;rft.genre=journal&amp;rft.issue=8&amp;rft.jtitle=iScience&amp;rft.pages=113184&amp;rft.pmc=12432456&amp;rft.pmid=40948565&amp;rft.volume=28.+Jahrgang" style="display:none">&nbsp;</span></span>
</li>
<li id="cite_note-Janouskovec2017-5"><span class="mw-cite-backlink"><a href="#cite_ref-Janouskovec2017_5-0">↑</a></span> <span class="reference-text">
Jan Janouškovec, Denis V. Tikhonenkov, Fabien Burki, Alexis T. Howe, Forest L. Rohwer, Alexander P. Mylnikov, Patrick J. Keeling: <i>A New Lineage of Eukaryotes Illuminates Early Mitochondrial Genome Reduction.</i> In: <i>Current Biology</i>, Band 27, Nr.&nbsp;23, 4. Dezember 2017, S.&nbsp;3717–3724; <a href="https://doi.org/10.1016/j.cub.2017.10.051" class="extiw external" title="doi:10.1016/j.cub.2017.10.051">doi:10.1016/j.cub.2017.10.051</a> (englisch). Dazu:
<ul><li>Katarina Zimmer; Denis Tikhonenkov (Bild): <a rel="nofollow" class="external text" href="https://www.the-scientist.com/the-nutshell/a-newly-identified-species-represents-its-own-eukaryotic-lineage-30601">A Newly Identified Species Represents Its Own Eukaryotic Lineage — The 10-micrometer-long flagellate cell might have a big story to tell about the evolution of eukaryotes</a>. Auf: The Scientist, 20. November 2017.</li></ul>
</span></li>
<li id="cite_note-Valt2025-6"><span class="mw-cite-backlink"><a href="#cite_ref-Valt2025_6-0">↑</a></span> <span class="reference-text">
Marek Valt, Tomáš Pánek, Seda Mirzoyan, Alexander K. Tice, Robert E. Jones, Vít Dohnálek, Pavel Doležal, Jiří Mikšátko, Johana Rotterová, Pavla Hrubá, Matthew W. Brown, Ivan Čepička: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Rare microbial relict sheds light on an ancient eukaryotic supergroup</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic"><a href="Nature" title="Nature">Nature</a></cite>. 19.&nbsp;November 2025, <a href="Internationale_Standardnummer_f%C3%BCr_fortlaufende_Sammelwerke" title="Internationale Standardnummer für fortlaufende Sammelwerke">ISSN</a>&nbsp;<span style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://zdb-katalog.de/list.xhtml?t=iss%3D%220028-0836%22&amp;key=cql">0028-0836</a></span>, <a href="ResearchGate" class="mw-redirect" title="ResearchGate">ResearchGate</a>:<a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/397765357">397765357</a>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1038/s41586-025-09750-0">10.1038/s41586-025-09750-0</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/41261123?dopt=Abstract">PMID 41261123</a> (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rfr_id=info:sid/de.wikipedia.org:Nebulidia&amp;rft.atitle=Rare+microbial+relict+sheds+light+on+an+ancient+eukaryotic+supergroup&amp;rft.au=Marek+Valt%2C+Tom%C3%A1%C5%A1+P%C3%A1nek%2C+Seda+Mirzoyan%2C+...&amp;rft.date=2025-11-19&amp;rft.doi=10.1038%2Fs41586-025-09750-0&amp;rft.genre=journal&amp;rft.issn=0028-0836&amp;rft.jtitle=Nature&amp;rft.pmid=41261123" style="display:none">&nbsp;</span> Dazu:
<ul><li>Vladimír Krylov: <a rel="nofollow" class="external text" href="https://natur.cuni.cz/en/news/9383-discovery-of-a-rare-protist-reveals-a-previously-unknown-branch-of-the-eukaryotic-tree-of-life">Discovery of a rare protist reveals a previously unknown branch of the eukaryotic tree of life</a>. <a href="Karls-Universit%C3%A4t" title="Karls-Universität">Karls-Universität</a>, Prag, 19. November 2025.</li></ul>
</span></li>
<li id="cite_note-Tikhonenkov2022-03-7"><span class="mw-cite-backlink"><a href="#cite_ref-Tikhonenkov2022-03_7-0">↑</a></span> <span class="reference-text">
Denis V. Tikhonenkov, Mahwash Jamy, Anastasia S. Borodina, Artem O. Belyaev, Dmitry G. Zagumyonnyi, Kristina I. Prokina, Alexander P. Mylnikov, Fabien Burki, Sergey A. Karpov: <i>On the origin of TSAR: morphology, diversity and phylogeny of Telonemia.</i> In: <i>Open Biology</i>, Band 12, Nr.&nbsp;3, März 2022, S.&nbsp;210325; <a href="https://doi.org/10.1098/rsob.210325" class="extiw external" title="doi:10.1098/rsob.210325">doi:10.1098/rsob.210325</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8924772/">PMC&nbsp;8924772</a> (freier Volltext), <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/35291881?dopt=Abstract">PMID 35291881</a>, Epub 16. März 2022 (englisch).</span>
</li>
<li id="cite_note-Tikhonenkov2023-8"><span class="mw-cite-backlink"><a href="#cite_ref-Tikhonenkov2023_8-0">↑</a></span> <span class="reference-text">
Denis V. Tikhonenkov, Kirill V. Mikhailov, Ryan M. R. Gawryluk, Patrick J. Keeling: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Provora</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic"><a href="Current_Biology" title="Current Biology">Current Biology</a></cite>. 33. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em">&nbsp;</span>15</span>, 7.&nbsp;August 2023, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em">&nbsp;</span>R790–R791</span>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1016/j.cub.2023.06.004">10.1016/j.cub.2023.06.004</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/37552939?dopt=Abstract">PMID 37552939</a> (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rfr_id=info:sid/de.wikipedia.org:Nebulidia&amp;rft.atitle=Provora&amp;rft.au=Denis+V.+Tikhonenkov%2C+Kirill+V.+Mikhailov%2C+Ryan+M.+R.+Gawryluk%2C+...&amp;rft.date=2023-08-07&amp;rft.doi=10.1016%2Fj.cub.2023.06.004&amp;rft.genre=journal&amp;rft.issue=15&amp;rft.jtitle=Current+Biology&amp;rft.pages=R790-R791&amp;rft.pmid=37552939&amp;rft.volume=33.+Jahrgang" style="display:none">&nbsp;</span> Siehe auch <a rel="nofollow" class="external text" href="https://www.cell.com/cms/10.1016/j.cub.2023.06.004/asset/ed01871e-a10f-4452-851a-d5059e809d66/main.assets/gr1_lrg.jpg">Graphical Abstract</a></span>
</li>
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