<!DOCTYPE html>
<html class="client-nojs vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-0 vector-toc-not-available vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-0 skin-theme-clientpref-day vector-sticky-header-enabled" lang="de" dir="ltr"><head>
<meta charset="UTF-8">
<title>Nebulidia</title>
<meta name="viewport" content="width=device-width, initial-scale=1.0">
<link rel="icon" type="image/png" href="./_res_/favicon.png">
<link rel="canonical" href="https://de.wikipedia.org/wiki/Nebulidia"> <link href="./_mw_/ext.cite.styles.css" rel="stylesheet" type="text/css">
<link href="./_mw_/ext.wikimediamessages.styles.css" rel="stylesheet" type="text/css">
<link href="./_mw_/skins.vector.icons.css" rel="stylesheet" type="text/css">
<link href="./_mw_/skins.vector.search.codex.styles.css" rel="stylesheet" type="text/css">
<link href="./_mw_/skins.vector.styles.css" rel="stylesheet" type="text/css">
<meta name="ResourceLoaderDynamicStyles" content="">
<link href="./_mw_/ext.gadget.citeRef.css" rel="stylesheet" type="text/css">
<link href="./_mw_/ext.gadget.defaultPlainlinks.css" rel="stylesheet" type="text/css">
<link href="./_mw_/ext.gadget.dewikiCommonHide.css" rel="stylesheet" type="text/css">
<link href="./_mw_/ext.gadget.dewikiCommonLayout.css" rel="stylesheet" type="text/css">
<link href="./_mw_/ext.gadget.dewikiCommonStyle.css" rel="stylesheet" type="text/css">
<link href="./_mw_/ext.gadget.dewikiDarkmode.css" rel="stylesheet" type="text/css">
<link href="./_mw_/ext.gadget.dewikiResponsive.css" rel="stylesheet" type="text/css">
<link href="./_mw_/ext.gadget.specialSearch.css" rel="stylesheet" type="text/css">
<link rel="stylesheet" type="text/css" href="./_mw_/site.styles.css">
<link rel="stylesheet" type="text/css" href="./_mw_/noscript.css">
<link rel="stylesheet" type="text/css" href="./_res_/footer.css">
<link rel="stylesheet" type="text/css" href="./_res_/vector-2022.css">
</head>
<body class="skin--responsive skin-vector skin-vector-search-vue mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject page-Nebulidia rootpage-Nebulidia skin-vector-2022 action-view">
<div class="mw-page-container">
<div class="mw-page-container-inner">
<div class="mw-content-container">
<main id="content" class="mw-body">
<header class="mw-body-header vector-page-titlebar">
<h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Nebulidia</span></h1>
</header>
<a id="top"></a>
<div id="bodyContent" class="vector-body ve-init-mw-desktopArticleTarget-targetContainer" aria-labelledby="firstHeading" data-mw-ve-target-container="">
<div id="contentSub">
<div id="mw-content-subtitle"></div>
</div>
<div id="mw-content-text" class="mw-body-content mw-content-ltr" lang="de" dir="ltr"><div class="mw-content-ltr mw-parser-output" lang="de" dir="ltr"><style data-mw-deduplicate="TemplateStyles:r183054365">
/* start https://de.wikipedia.org/ */
.mw-parser-output .Person{font-variant:small-caps}.mw-parser-output table.taxobox{background:white;border:1px solid #72777d;border-collapse:collapse;clear:right;float:right;margin:1em 0 1em 1em}.mw-parser-output table.taxobox>*>*>th{background:#9bcd9b;border:1px solid #72777d;text-align:center}.mw-parser-output table.taxobox.palaeobox>*>*>th{background:#e7dcc3}.mw-parser-output table.taxobox.parabox>*>*>th{background:#b9b9b9}.mw-parser-output table.taxobox>*>*>td.Person,.mw-parser-output table.taxobox>*>*>td.taxo-name,.mw-parser-output table.taxobox>*>*>td.taxo-bild,.mw-parser-output table.taxobox>*>*>td.taxo-zeit{text-align:center}
/* end https://de.wikipedia.org/ */
</style>
<table cellpadding="2" class="float-right taxobox infobox" id="Vorlage_Taxobox" style="width:300px;margin-top:0.5em;" summary="Taxobox">
<tbody><tr>
<th>Nebulidae
</th></tr>
<tr>
<th><a href="Systematik_(Biologie)" title="Systematik (Biologie)">Systematik</a>
</th></tr>
<tr>
<td>
<table class="toptextcells" style="width:100%;">
<tbody><tr>
<td><i><a href="Reich_(Biologie)" title="Reich (Biologie)">Reich</a>:</i>
</td>
<td><a href="Disparia" title="Disparia">Disparia</a>
</td></tr>
<tr>
<td><i>ohne Rang:</i>
</td>
<td><a href="Provora" title="Provora">Provora</a>
</td></tr>
<tr>
<td><i><a href="Stamm_(Biologie)" title="Stamm (Biologie)">Stamm</a>:</i>
</td>
<td>Nebulidia
</td></tr>
<tr>
<td><i><a href="Klasse_(Biologie)" title="Klasse (Biologie)">Klasse</a>:</i>
</td>
<td>Nebulidea
</td></tr>
<tr>
<td><i><a href="Ordnung_(Biologie)" title="Ordnung (Biologie)">Ordnung</a>:</i>
</td>
<td>Nebulidida
</td></tr>
<tr>
<td><i><a href="Familie_(Biologie)" title="Familie (Biologie)">Familie</a>:</i>
</td>
<td>Nebulidae
</td></tr>
</tbody></table>
</td></tr>
<tr>
<th><a href="Nomenklatur_(Biologie)" title="Nomenklatur (Biologie)">Wissenschaftlicher Name</a> des <a href="Stamm_(Biologie)" title="Stamm (Biologie)">Stamms</a>
</th></tr>
<tr>
<td class="taxo-name">Nebulidia
</td></tr>
<tr>
<td class="Person">Tikhonenkov <i>et al.</i> 2022<sup id="cite_ref-Tikhonenkov2022-12_1-0" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
</td></tr>
<tr>
<th><a href="Nomenklatur_(Biologie)" title="Nomenklatur (Biologie)">Wissenschaftlicher Name</a> der <a href="Klasse_(Biologie)" title="Klasse (Biologie)">Klasse</a>
</th></tr>
<tr>
<td class="taxo-name">Nebulidea
</td></tr>
<tr>
<td class="Person">Tikhonenkov <i>et al.</i> 2022<sup id="cite_ref-Tikhonenkov2022-12_1-1" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
</td></tr>
<tr>
<th><a href="Nomenklatur_(Biologie)" title="Nomenklatur (Biologie)">Wissenschaftlicher Name</a> der <a href="Ordnung_(Biologie)" title="Ordnung (Biologie)">Ordnung</a>
</th></tr>
<tr>
<td class="taxo-name">Nebulidida
</td></tr>
<tr>
<td class="Person">Tikhonenkov <i>et al.</i> 2022<sup id="cite_ref-Tikhonenkov2022-12_1-2" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
</td></tr>
<tr>
<th><a href="Nomenklatur_(Biologie)" title="Nomenklatur (Biologie)">Wissenschaftlicher Name</a> der <a href="Familie_(Biologie)" title="Familie (Biologie)">Familie</a>
</th></tr>
<tr>
<td class="taxo-name">Nebulidae
</td></tr>
<tr>
<td class="Person">Tikhonenkov <i>et al.</i> 2022<sup id="cite_ref-Tikhonenkov2022-12_1-3" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
</td></tr>
</tbody></table>
<p><b>Nebulidia</b> ist ein <a href="Phylum" class="mw-redirect" title="Phylum">Phylum</a> (<a href="Stamm_(Biologie)" title="Stamm (Biologie)">Stamm</a>) räuberischer <a href="Mikrobiell" class="mw-redirect" title="Mikrobiell">mikrobieller</a> <a href="Eukaryoten" title="Eukaryoten">Eukaryoten</a> (<a href="Mikroeukaryoten" class="mw-redirect" title="Mikroeukaryoten">Mikroeukaryoten</a>, <a href="Protisten" title="Protisten">Protisten</a>) in der Supergruppe <a href="Provora" title="Provora">Provora</a>. Das Phylum dieser <a href="Einzeller" title="Einzeller">Einzeller</a> umfasst <a href="Monotypisch" title="Monotypisch">monotypisch</a> nur die <a href="Klasse_(Biologie)" title="Klasse (Biologie)">Klasse</a> <b>Nebulidea</b>, mit der einzigen <a href="Ordnung_(Biologie)" title="Ordnung (Biologie)">Ordnung</a> <b>Nebulidida</b>, und diese die eine <a href="Familie_(Biologie)" title="Familie (Biologie)">Familie</a> <b>Nebulidae</b> – diese Gruppe (<a href="Klade" title="Klade">Klade</a>) wird informell auch als ‚Nebuliden‘ (<span style="font-style:normal;font-weight:normal"><a href="Englische_Sprache" title="Englische Sprache">englisch</a></span> <span lang="en-Latn" style="font-style:italic">nebulids</span>) bezeichnet. Die Familie Nebulidae beinhaltet die <a href="Gattung_(Biologie)" title="Gattung (Biologie)">Gattungen</a> <i><a href="Nebulomonas" class="mw-redirect" title="Nebulomonas">Nebulomonas</a></i> und <i><a href="Ancoracysta" class="mw-redirect" title="Ancoracysta">Ancoracysta</a></i>.<sup id="cite_ref-NCBI_2-0" class="reference"><a href="#cite_note-NCBI-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Tikhonenkov2022-12_1-4" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Belyaev2024_3-0" class="reference"><a href="#cite_note-Belyaev2024-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup>
</p>
<div class="mw-heading mw-heading2"><h2 id="Nahrungserwerb">Nahrungserwerb</h2></div>
<p>Zum Nahrungserwerb besitzen die Nebulidia <a href="Organell#Taxonspezifische_Organellen" title="Organell">spezielle Organellen</a>, dei bei der Gattung <i><a href="Ancoracysta" class="mw-redirect" title="Ancoracysta">Ancoracysta</a></i> Ancoracysten und bei der Gattung <i><a href="Nebulomonas" class="mw-redirect" title="Nebulomonas">Nebulomonas</a></i> Amphoracysten (Toxicysten) genannt werden.<sup id="cite_ref-Tikhonenkov2022-12_1-5" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>
</p>
<div class="mw-heading mw-heading2"><h2 id="Systematik">Systematik</h2></div>
<p>Die Systematik der Klade lautet mit Stand 9. Dezember 2025 wie folgt:
</p><p><a href="Reich_(Biologie)" title="Reich (Biologie)">Reich</a> <a href="Disparia" title="Disparia">Disparia</a> <small><style data-mw-deduplicate="TemplateStyles:r261921330">
/* start https://de.wikipedia.org/ */
.mw-parser-output .Person{font-variant:small-caps}
/* end https://de.wikipedia.org/ */
</style><span class="Person h-card">Valt & Čepička</span> 2025</small><sup>(V)</sup> <span style="white-space:nowrap">[Promethea <small><span class="Person h-card">Zlatogursky, Boscaro & Keeling</span>, 2025</small><sup id="cite_ref-Zlatogursky2025_4-0" class="reference"><a href="#cite_note-Zlatogursky2025-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup>]</span>
</p>
<ul><li><a href="Klade" title="Klade">Klade</a> <a href="Membrifera" title="Membrifera">Membrifera</a> <small><span class="Person h-card">Valt & Čepička</span> 2025</small><sup>(V)</sup>
<ul><li><a href="Phylum" class="mw-redirect" title="Phylum">Phylum</a> <a href="Caelestes" title="Caelestes">Caelestes</a> <small><span class="Person h-card">Valt & Čepička</span> 2025</small><sup>(V)</sup></li>
<li>Phylum <a href="Hemimastigophora" title="Hemimastigophora">Hemimastigophora</a> <small><span class="Person h-card"><a href="Wilhelm_Foissner" title="Wilhelm Foissner">W. Foissner</a>, H. Blatterer & I. Foissner</span>, 1988</small></li></ul></li>
<li>Klade <a href="Provora" title="Provora">Provora</a> <small><span class="Person h-card">Tikhonenkov</span> <i>et al.</i> 2022</small><sup>(N,T,V)</sup>
<ul><li>Phylum <b>Nebulidia</b> <small><span class="Person h-card">Tikhonenkov</span> <i>et al.</i> 2022</small><sup>(N,T)</sup>
<ul><li>Klasse <b>Nebulidea</b><sup>(N)</sup> <small><span class="Person h-card">Tikhonenkov</span> <i>et al.</i> 2022</small><sup>(T)</sup>
<ul><li>Ordnung <b>Nebulidida</b><sup>(N)</sup> <small><span class="Person h-card">Tikhonenkov</span> <i>et al.</i> 2022</small><sup>(T)</sup>
<ul><li>Familie <b>Nebulidae</b><sup>(N)</sup> <small><span class="Person h-card">Tikhonenkov</span> <i>et al.</i> 2022</small><sup>(T)</sup>
<ul><li>Gattung <i><a href="Ancoracysta" class="mw-redirect" title="Ancoracysta">Ancoracysta</a></i> <small><span class="Person h-card">Janouškovec, Tikhonenkov, Burki, Howe, Rohwer, Mylnikov & Keeling</span> 2017</small><sup>(N)</sup><sup id="cite_ref-Janouskovec2017_5-0" class="reference"><a href="#cite_note-Janouskovec2017-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li>
<li>Gattung <i><b><a href="Nebulomonas" class="mw-redirect" title="Nebulomonas">Nebulomonas</a></b></i> <small><span class="Person h-card">Tikhonenkov</span> <i>et al.</i> 2022</small><sup>(N,T)</sup></li></ul></li></ul></li></ul></li></ul></li>
<li>Phylum <a href="Nibbleridia" title="Nibbleridia">Nibbleridia</a> <small><span class="Person h-card">Tikhonenkov</span> <i>et al.</i> 2022</small><sup>(N,T)</sup><br>– mit Gattungen <i><a href="Nibbleromonas" title="Nibbleromonas">Nibbleromonas</a></i>, <i><a href="Ubysseya" class="mw-redirect" title="Ubysseya">Ubysseya</a></i></li></ul></li></ul>
<hr>
<dl><dd>(N) – <span lang="en"><a href="National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">National Center for Biotechnology Information</a></span> (NCBI)<sup id="cite_ref-NCBI_2-1" class="reference"><a href="#cite_note-NCBI-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></dd>
<dd>(V) – Marek Valt, Alexander K. Tice, Ivan Čepička <i>et al.</i> (2025)<sup id="cite_ref-Valt2025_6-0" class="reference"><a href="#cite_note-Valt2025-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup></dd>
<dd>(T) – Tikhonenkov <i>et al.</i><sup id="cite_ref-Tikhonenkov2022-03_7-0" class="reference"><a href="#cite_note-Tikhonenkov2022-03-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Tikhonenkov2022-12_1-6" class="reference"><a href="#cite_note-Tikhonenkov2022-12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Tikhonenkov2023_8-0" class="reference"><a href="#cite_note-Tikhonenkov2023-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></dd></dl>
<div class="mw-heading mw-heading2"><h2 id="Phylogenie">Phylogenie</h2></div>
<p>Die phylogenetischen Beziehungen zwischen den beschriebenen Arten der Nebulidia werden wie folgt dargestellt:<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>1.1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>3.1<span class="cite-bracket">]</span></a></sup>
</p>
<table class="klade" style="border-collapse:collapse;line-height:100%;font-size:100%;">
<tbody><tr>
<td style="min-width:1em; border-bottom:2px solid; vertical-align:bottom; text-align:center;"> <a href="Provora" title="Provora">Provora</a><sup><sup><span style="visibility:hidden;">.</span></sup></sup>
</td>
<td rowspan="2" style="padding-bottom:3px;">
<table class="klade" style="border-collapse:collapse;line-height:100%;">
<tbody><tr>
<td style="min-width:1em; border-bottom:2px solid; vertical-align:bottom; text-align:center;"> <b>Nebulidia</b><sup><span style="visibility:hidden;">.</span></sup>
</td>
<td rowspan="2" style="padding-bottom:3px;">
<table class="klade" style="border-collapse:collapse;line-height:100%;">
<tbody><tr>
<td style="min-width:1em; border-bottom:2px solid; vertical-align:bottom; text-align:center;"> <i>Nebulomonas</i>
</td>
<td rowspan="2" style="padding-bottom:3px;padding-left:0.5em;">
<p><i><a href="Nebulomonas_marisrubri" title="Nebulomonas marisrubri">N. marisrubri</a></i>
</p>
</td></tr>
<tr>
<td style="border-left:2px solid; vertical-align:top;"><br>
</td></tr>
<tr>
<td style="border-left:2px solid; border-bottom:2px solid; vertical-align:bottom; text-align:center;"> <i>Ancoracysta</i>
</td>
<td rowspan="2" style="padding-bottom:3px;padding-left:0.5em;">
<p><i><a href="Ancoracysta_twista" title="Ancoracysta twista">A. twista</a></i>
</p>
</td></tr>
<tr>
<td valign="top"><br>
</td></tr></tbody></table><style data-mw-deduplicate="TemplateStyles:r257636185">
/* start https://de.wikipedia.org/ */
.mw-parser-output .tptable{border-collapse:collapse;font-size:90%;margin:1em 0;max-width:100%;text-align:center;box-sizing:border-box;line-height:130%}.mw-parser-output .tptblxl{border-collapse:collapse;font-size:90%;margin:1em 0;max-width:100%;text-align:center;box-sizing:border-box;line-height:1.5em}.mw-parser-output .tpheader{border:1px solid #999999}.mw-parser-output .tpcaptiont{text-align:center;font-size:90%;vertical-align:top}.mw-parser-output .tpcaptionm{text-align:center;font-size:90%;vertical-align:middle}.mw-parser-output .tpcaptionb{text-align:center;font-size:90%;vertical-align:bottom}.mw-parser-output .tpseed{border:1px solid #999999;background-color:#6C7A93;color:#FFFFFF;height:20px}.mw-parser-output .tpteam{border:1px solid #999999;padding:0 0.5em;text-align:left;height:20px}.mw-parser-output .tpscore{border:1px solid #999999;height:20px}.mw-parser-output .tpgold{background:gold}.mw-parser-output .tpsilver{background:silver}.mw-parser-output .tpbronze{background:#CC9966}.mw-parser-output .tpgroup{text-align:center}.mw-parser-output .tpeditonly{font-size:1px;color:transparent;width:3px}.mw-parser-output .tp0000{border-style:solid;border-width:0 0 0 0}.mw-parser-output .tp0002{border-style:solid;border-width:0 0 0 2px}.mw-parser-output .tp0020{border-style:solid;border-width:0 0 2px 0}.mw-parser-output .tp0022{border-style:solid;border-width:0 0 2px 2px}.mw-parser-output .tp0200{border-style:solid;border-width:0 2px 0 0}.mw-parser-output .tp0202{border-style:solid;border-width:0 2px 0 2px}.mw-parser-output .tp0220{border-style:solid;border-width:0 2px 2px 0}.mw-parser-output .tp0222{border-style:solid;border-width:0 2px 2px 2px}.mw-parser-output .tp2000{border-style:solid;border-width:2px 0 0 0}.mw-parser-output .tp2002{border-style:solid;border-width:2px 0 0 2px}.mw-parser-output .tp2020{border-style:solid;border-width:2px 0 2px 0}.mw-parser-output .tp2022{border-style:solid;border-width:2px 0 2px 2px}.mw-parser-output .tp2200{border-style:solid;border-width:2px 2px 0 0}.mw-parser-output .tp2202{border-style:solid;border-width:2px 2px 0 2px}.mw-parser-output .tp2220{border-style:solid;border-width:2px 2px 2px 0}.mw-parser-output .tpbb{border-bottom:2px solid #777777}.mw-parser-output .tptt{border-top:2px solid #777777}.mw-parser-output .tpbox{border-collapse:collapse;font-size:90%;margin:0;text-align:center;box-sizing:border-box;line-height:120%}.mw-parser-output .tpeditonly{display:none}.mw-parser-output .tpoption{background:#ff9900;margin:0;padding:1px;font-style:italic}@media screen{html.skin-theme-clientpref-night .mw-parser-output .tpseed{border:1px solid #DDDDDD;background:#B6BCC9;color:#000000;height:20px}html.skin-theme-clientpref-night .mw-parser-output .tptable td{border-color:#DDDDDD}html.skin-theme-clientpref-night .mw-parser-output .tpbb{border-bottom:2px solid #DDDDDD}html.skin-theme-clientpref-night .mw-parser-output .tptt{border-top:2px solid #DDDDDD}}
/* end https://de.wikipedia.org/ */
</style>
</td></tr>
<tr>
<td style="border-left:2px solid; vertical-align:top;"><br>
</td></tr>
<tr>
<td style="border-left:2px solid; border-bottom:2px solid; vertical-align:bottom; text-align:center;"> <a href="Nibbleridia" title="Nibbleridia">Nibbleridia</a>
</td>
<td rowspan="2" style="padding-bottom:3px;padding-left:0.5em;">
<p><i><a href="Ubysseya_fretuma" title="Ubysseya fretuma">Ubysseya fretuma</a></i>, <i><a href="Nibbleromonas" title="Nibbleromonas">Nibbleromonas spp.</a></i>
</p>
</td></tr>
<tr>
<td valign="top"><br>
</td></tr></tbody></table>
</td></tr>
<tr>
<td valign="top"><br><span style="display:none;">Vorlage:Klade/Wartung/Style</span>
</td></tr></tbody></table>
<div class="mw-heading mw-heading2"><h2 id="Etymologie">Etymologie</h2></div>
<p>Die Bezeichnungen leiten sich ab von <span style="font-style:normal;font-weight:normal"><a href="Latein" title="Latein">lateinisch</a></span> <span lang="la-Latn" style="font-style:italic">nebulosus</span>, deutsch <span lang="de" style="font-style:normal;font-weight:normal">‚neblig‘</span>, <a href="Englische_Sprache" title="Englische Sprache">englisch</a> <span lang="en" style="font-style:italic;font-weight:normal">misty</span>, der jeweilige Suffix kennzeichnet die taxonomische Rangstufe.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>1.2<span class="cite-bracket">]</span></a></sup>
</p>
<div class="mw-heading mw-heading2"><h2 id="Weiterführende_Literatur"><span id="Weiterf.C3.BChrende_Literatur"></span>Weiterführende Literatur</h2></div>
<ul><li>Sei Suzuki-Tellier, Federica Miano, Seyed Saeed Asadzadeh, Alastair G. B. Simpson, Thomas Kiørboe: <i>Foraging mechanisms in excavate flagellates shed light on the functional ecology of early eukaryotes.</i> In: <i><a href="PNAS" class="mw-redirect" title="PNAS">PNAS</a></i>, Band 121, Nr. 22, 23. Mai 2024, S. e2317264121; <a href="https://doi.org/10.1073/pnas.2317264121" class="extiw external" title="doi:10.1073/pnas.2317264121">doi:10.1073/pnas.2317264121</a>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/38781211?dopt=Abstract">PMID 38781211</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11145212/">PMC 11145212</a> (freier Volltext), <a href="ResearchGate" class="mw-redirect" title="ResearchGate">ResearchGate</a>:<a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/375822320">375822320</a> (englisch).</li>
<li>Łukasz Lamża: <i>Deep-branching eukaryotes and early events in protist evolution.</i> In: <i>Biological Reviews</i>, 9. November 2025; <a href="https://doi.org/10.1111/brv.70101" class="extiw external" title="doi:10.1111/brv.70101">doi:10.1111/brv.70101</a> (englisch).</li></ul>
<div class="mw-heading mw-heading2"><h2 id="Einzelnachweise">Einzelnachweise</h2></div>
<ol class="references">
<li id="cite_note-Tikhonenkov2022-12-1"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-Tikhonenkov2022-12_1-0">a</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-1">b</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-2">c</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-3">d</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-4">e</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-5">f</a></sup> <sup><a href="#cite_ref-Tikhonenkov2022-12_1-6">g</a></sup></span> <span class="reference-text">
Denis Victorovich Tikhonenkov, Kirill V. Mikhailov, Ryan M. R. Gawryluk, Artem O. Belyaev, Varsha Mathur, Sergey A. Karpov, Dmitry G. Zagumyonnyi, Anastasia S. Borodina, Kristina I. Prokina, Alexander P. Mylnikov, Vladimir V. Aleoshin, Patrick J. Keeling: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Microbial predators form a new supergroup of eukaryotes</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic"><a href="Nature" title="Nature">Nature</a></cite>. 612. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>7941</span>, 7. Dezember 2022, <a href="Internationale_Standardnummer_f%C3%BCr_fortlaufende_Sammelwerke" title="Internationale Standardnummer für fortlaufende Sammelwerke">ISSN</a> <span style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://zdb-katalog.de/list.xhtml?t=iss%3D%220028-0836%22&key=cql">0028-0836</a></span>, <a href="ResearchGate" class="mw-redirect" title="ResearchGate">ResearchGate</a>:<a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/366093658">366093658</a>, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>714–719</span>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1038/s41586-022-05511-5">10.1038/s41586-022-05511-5</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/36477531?dopt=Abstract">PMID 36477531</a>, <a href="Bibcode" title="Bibcode">bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2022Natur.612..714T">2022Natur.612..714T</a> (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:Nebulidia&rft.atitle=Microbial+predators+form+a+new+supergroup+of+eukaryotes&rft.au=Denis+Victorovich+Tikhonenkov%2C+Kirill+V.+Mikhailov%2C+Ryan+M.+R.+Gawryluk%2C+...&rft.date=2022-12-07&rft.doi=10.1038%2Fs41586-022-05511-5&rft.genre=journal&rft.issn=0028-0836&rft.issue=7941&rft.jtitle=Nature&rft.pages=714-719&rft.pmid=36477531&rft.volume=612.+Jahrgang" style="display:none"> </span> Dazu:
<dl><dd><dl><dd><ul><li><a rel="nofollow" class="external text" href="https://static-content.springer.com/esm/art%3A10.1038%2Fs41586-022-05511-5/MediaObjects/41586_2022_5511_MOESM1_ESM.pdf">Supplementay Information</a> (PDF; 0,5 MB)</li>
<li>Nadja Podbregar: <a rel="nofollow" class="external text" href="https://www.scinexx.de/news/biowissen/neue-supergruppe-am-baum-des-lebens/">Neue Supergruppe am Baum des Lebens</a> – Uralte Gruppe räuberischer Einzeller unterscheidet sich von allen anderen Lebewesen. Auf: <a href="Scinexx.de" class="mw-redirect" title="Scinexx.de">scinexx.de</a> vom 15. Dezember 2022.</li>
<li><a rel="nofollow" class="external text" href="https://www.sci.news/biology/provora-11460.html">Meet Provora, New Supergroup of Eukaryotes</a>. Auf <a href="Sci.News" title="Sci.News">Sci.News</a> vom 8. Dezember 2022.</li>
<li>Stefan Labbé: <a rel="nofollow" class="external text" href="https://www.vancouverisawesome.com/highlights/nibblers-and-lions-bc-researchers-discover-new-supergroup-on-tree-of-life-6220625">'Nibblers and Lions': B.C. researchers discover new 'supergroup' on tree of life</a>. Auf: Vancouver is Awesome vom 8. Dezember 2022.</li></ul></dd></dl></dd></dl>
<ol class="mw-subreference-list"><li id="cite_note-9"><span class="mw-cite-backlink"><a href="#cite_ref-9">↑</a></span> <span class="reference-text">S. 717</span>
</li>
<li id="cite_note-11"><span class="mw-cite-backlink"><a href="#cite_ref-11">↑</a></span> <span class="reference-text">Suppl. Inf.</span>
</li>
</ol></span></li>
<li id="cite_note-NCBI-2"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-NCBI_2-0">a</a></sup> <sup><a href="#cite_ref-NCBI_2-1">b</a></sup></span> <span class="reference-text">
<a href="National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a> Taxonomy Browser: <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?command=show&mode=tree&id=3004206&lvl=3">Provora</a>, Details: <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?command=show&mode=node&id=3004206">Provora in [Tikhonenkov DV et al. (2022)]</a>. Graphisch: <a rel="nofollow" class="external text" href="https://lifemap.cnrs.fr/tree?efficiency-mode=true&tool=search&tid=3004206">Provora</a>, auf <a href="Lifemap" class="mw-redirect" title="Lifemap">Lifemap</a>.</span>
</li>
<li id="cite_note-Belyaev2024-3"><span class="mw-cite-backlink"><a href="#cite_ref-Belyaev2024_3-0">↑</a></span> <span class="reference-text">
Artem O. Belyaev, Sergei A. Karpov, Patrick J. Keeling, Denis V. Tikhonenkov: <cite class="lang" lang="en" dir="auto" style="font-style:italic">The nature of 'jaws': a new predatory representative of Provora and the ultrastructure of nibbling protists</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Open Biology</cite>. 14. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>12</span>, 18. Dezember 2024, <a href="ResearchGate" class="mw-redirect" title="ResearchGate">ResearchGate</a>:<a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/387173990">387173990</a>, <a rel="nofollow" class="external text" href="https://www3.botany.ubc.ca/keeling/PDF/24jaws.pdf">PDF</a> (botany.ubc.ca), <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>240158</span>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1098/rsob.240158">10.1098/rsob.240158</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/39689855?dopt=Abstract">PMID 39689855</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11651884/">PMC 11651884</a> (freier Volltext) – (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:Nebulidia&rft.atitle=The+nature+of+%27jaws%27%3A+a+new+predatory+representative+of+Provora+and+the+ultrastructure+of+nibbling+protists&rft.au=Artem+O.+Belyaev%2C+Sergei+A.+Karpov%2C+Patrick+J.+Keeling%2C+...&rft.date=2024-12-18&rft.doi=10.1098%2Frsob.240158&rft.genre=journal&rft.issue=12&rft.jtitle=Open+Biology&rft.pages=240158&rft.pmc=11651884&rft.pmid=39689855&rft.volume=14.+Jahrgang" style="display:none"> </span></span>
<ol class="mw-subreference-list"><li id="cite_note-10"><span class="mw-cite-backlink"><a href="#cite_ref-10">↑</a></span> <span class="reference-text">S. 11-12</span>
</li>
</ol></li>
<li id="cite_note-Zlatogursky2025-4"><span class="mw-cite-backlink"><a href="#cite_ref-Zlatogursky2025_4-0">↑</a></span> <span class="reference-text">
Vasily Zlatogursky, Vittorio Boscaro, Gordon Lax, Matias Wanntorp, Nina Pohl, Fabien Burki, Patrick J. Keeling: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Phylogenetic position and mitochondrial genome evolution of “orphan” eukaryotic lineages</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic">iScience</cite>. 28. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>8</span>, August 2025, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>113184</span>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1016/j.isci.2025.113184">10.1016/j.isci.2025.113184</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/40948565?dopt=Abstract">PMID 40948565</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC12432456/">PMC 12432456</a> (freier Volltext) – (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:Nebulidia&rft.atitle=Phylogenetic+position+and+mitochondrial+genome+evolution+of+%E2%80%9Corphan%E2%80%9D+eukaryotic+lineages&rft.au=Vasily+Zlatogursky%2C+Vittorio+Boscaro%2C+Gordon+Lax%2C+...&rft.date=2025-08&rft.doi=10.1016%2Fj.isci.2025.113184&rft.genre=journal&rft.issue=8&rft.jtitle=iScience&rft.pages=113184&rft.pmc=12432456&rft.pmid=40948565&rft.volume=28.+Jahrgang" style="display:none"> </span></span>
</li>
<li id="cite_note-Janouskovec2017-5"><span class="mw-cite-backlink"><a href="#cite_ref-Janouskovec2017_5-0">↑</a></span> <span class="reference-text">
Jan Janouškovec, Denis V. Tikhonenkov, Fabien Burki, Alexis T. Howe, Forest L. Rohwer, Alexander P. Mylnikov, Patrick J. Keeling: <i>A New Lineage of Eukaryotes Illuminates Early Mitochondrial Genome Reduction.</i> In: <i>Current Biology</i>, Band 27, Nr. 23, 4. Dezember 2017, S. 3717–3724; <a href="https://doi.org/10.1016/j.cub.2017.10.051" class="extiw external" title="doi:10.1016/j.cub.2017.10.051">doi:10.1016/j.cub.2017.10.051</a> (englisch). Dazu:
<ul><li>Katarina Zimmer; Denis Tikhonenkov (Bild): <a rel="nofollow" class="external text" href="https://www.the-scientist.com/the-nutshell/a-newly-identified-species-represents-its-own-eukaryotic-lineage-30601">A Newly Identified Species Represents Its Own Eukaryotic Lineage — The 10-micrometer-long flagellate cell might have a big story to tell about the evolution of eukaryotes</a>. Auf: The Scientist, 20. November 2017.</li></ul>
</span></li>
<li id="cite_note-Valt2025-6"><span class="mw-cite-backlink"><a href="#cite_ref-Valt2025_6-0">↑</a></span> <span class="reference-text">
Marek Valt, Tomáš Pánek, Seda Mirzoyan, Alexander K. Tice, Robert E. Jones, Vít Dohnálek, Pavel Doležal, Jiří Mikšátko, Johana Rotterová, Pavla Hrubá, Matthew W. Brown, Ivan Čepička: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Rare microbial relict sheds light on an ancient eukaryotic supergroup</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic"><a href="Nature" title="Nature">Nature</a></cite>. 19. November 2025, <a href="Internationale_Standardnummer_f%C3%BCr_fortlaufende_Sammelwerke" title="Internationale Standardnummer für fortlaufende Sammelwerke">ISSN</a> <span style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://zdb-katalog.de/list.xhtml?t=iss%3D%220028-0836%22&key=cql">0028-0836</a></span>, <a href="ResearchGate" class="mw-redirect" title="ResearchGate">ResearchGate</a>:<a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/397765357">397765357</a>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1038/s41586-025-09750-0">10.1038/s41586-025-09750-0</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/41261123?dopt=Abstract">PMID 41261123</a> (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:Nebulidia&rft.atitle=Rare+microbial+relict+sheds+light+on+an+ancient+eukaryotic+supergroup&rft.au=Marek+Valt%2C+Tom%C3%A1%C5%A1+P%C3%A1nek%2C+Seda+Mirzoyan%2C+...&rft.date=2025-11-19&rft.doi=10.1038%2Fs41586-025-09750-0&rft.genre=journal&rft.issn=0028-0836&rft.jtitle=Nature&rft.pmid=41261123" style="display:none"> </span> Dazu:
<ul><li>Vladimír Krylov: <a rel="nofollow" class="external text" href="https://natur.cuni.cz/en/news/9383-discovery-of-a-rare-protist-reveals-a-previously-unknown-branch-of-the-eukaryotic-tree-of-life">Discovery of a rare protist reveals a previously unknown branch of the eukaryotic tree of life</a>. <a href="Karls-Universit%C3%A4t" title="Karls-Universität">Karls-Universität</a>, Prag, 19. November 2025.</li></ul>
</span></li>
<li id="cite_note-Tikhonenkov2022-03-7"><span class="mw-cite-backlink"><a href="#cite_ref-Tikhonenkov2022-03_7-0">↑</a></span> <span class="reference-text">
Denis V. Tikhonenkov, Mahwash Jamy, Anastasia S. Borodina, Artem O. Belyaev, Dmitry G. Zagumyonnyi, Kristina I. Prokina, Alexander P. Mylnikov, Fabien Burki, Sergey A. Karpov: <i>On the origin of TSAR: morphology, diversity and phylogeny of Telonemia.</i> In: <i>Open Biology</i>, Band 12, Nr. 3, März 2022, S. 210325; <a href="https://doi.org/10.1098/rsob.210325" class="extiw external" title="doi:10.1098/rsob.210325">doi:10.1098/rsob.210325</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8924772/">PMC 8924772</a> (freier Volltext), <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/35291881?dopt=Abstract">PMID 35291881</a>, Epub 16. März 2022 (englisch).</span>
</li>
<li id="cite_note-Tikhonenkov2023-8"><span class="mw-cite-backlink"><a href="#cite_ref-Tikhonenkov2023_8-0">↑</a></span> <span class="reference-text">
Denis V. Tikhonenkov, Kirill V. Mikhailov, Ryan M. R. Gawryluk, Patrick J. Keeling: <cite class="lang" lang="en" dir="auto" style="font-style:italic">Provora</cite>. In: <cite class="lang" lang="en" dir="auto" style="font-style:italic"><a href="Current_Biology" title="Current Biology">Current Biology</a></cite>. 33. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>15</span>, 7. August 2023, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>R790–R791</span>, <a href="Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1016/j.cub.2023.06.004">10.1016/j.cub.2023.06.004</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="https://www.ncbi.nlm.nih.gov/pubmed/37552939?dopt=Abstract">PMID 37552939</a> (englisch).<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:Nebulidia&rft.atitle=Provora&rft.au=Denis+V.+Tikhonenkov%2C+Kirill+V.+Mikhailov%2C+Ryan+M.+R.+Gawryluk%2C+...&rft.date=2023-08-07&rft.doi=10.1016%2Fj.cub.2023.06.004&rft.genre=journal&rft.issue=15&rft.jtitle=Current+Biology&rft.pages=R790-R791&rft.pmid=37552939&rft.volume=33.+Jahrgang" style="display:none"> </span> Siehe auch <a rel="nofollow" class="external text" href="https://www.cell.com/cms/10.1016/j.cub.2023.06.004/asset/ed01871e-a10f-4452-851a-d5059e809d66/main.assets/gr1_lrg.jpg">Graphical Abstract</a></span>
</li>
</ol></div><!--htdig_noindex--><div><div class="zim-footer">
Dieser Artikel wurde von <a class="external text" title="Zuletzt bearbeitet am 2025-12-26" href="https://de.wikipedia.org/wiki/?title=Nebulidia&oldid=262743963">Wikipedia</a> herausgegeben. Der Text ist unter <a class="external text" href="https://creativecommons.org/licenses/by-sa/4.0/deed.de">Creative Commons Attribution-Share Alike 4.0</a> verfügbar, sofern nicht anders angegeben. Für die Mediendateien können zusätzliche Bedingungen gelten.
</div>
</div><!--/htdig_noindex--></div>
</div>
</main>
</div>
</div>
</div>
<script src="./_webp_/webpHandler.js"></script>
</body></html>